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It was a different story offscreen: The couple managed to date quietly for the first two years of the show, with no one the wiser until they broke up.
The 'Big Bang Theory' earned its first Emmy nominations in 2009 following its second season.
The trio posed with then-Sony Pictures boss Amy Pascal at the movie's January 2015 premiere in Los Angeles.
By the spring of 2016, Cuoco, a longtime horsewoman, was dating professional equestrian Karl Cook.
(From left: Amy Davidson, Katey Sagal, James Garner, Martin Spanjers and Cuoco.) On Sept.
24, 2007, Cuoco debuted as Penny, the female lead in CBS' 'The Big Bang Theory.' Her character, the next-door neighbor of physicists Sheldon Cooper (Jim Parsons) and Leonard Hofstadter (Johnny Galecki), was not in the original pilot.
Since they began dating in 2016, Cuoco, a fellow rider and jumper, has popped up at equestrian competitions.
In April, she accompanied Cook to an FEI World Cup event, marking her first-first ever visit to Nebraska, the home state of her In 2000, Cuoco, right, played 'Brady Bunch' actress Maureen Mc Cormick in 2000's 'Growing Up Brady,' a TV movie about life behind the scenes of the classic sitcom. C.', Adam Brody, as Barry Williams and Caitlin Cullum as Eve Plumb.
Cuoco was cast as Bridget "Beach" Hennessy, the eldest daughter on the ABC sitcom '8 Simple Rules,' which starred John Ritter and Katey Sagal.
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A source confirms to PEOPLE that Mel Gibson has struck up a romance with a young film writer who also happens to be a champion equestrian vaulter.
Here, Kaley Cuoco and co-stars Jim Parsons and Johnny Galecki appear onstage as presenters at that year's telecast.
The more time Penny spent around the boys on 'Big Bang,' the more science and nerd trivia knowledge she began to accumulate.